WOODSIDE (CBSLA/AP) — Koko, the gorilla who mastered sign language, has died.

The Gorilla Foundation says the 46-year-old western lowland gorilla died in her sleep at the foundation’s preserve in California’s Santa Cruz mountains on Tuesday.

Koko was born at the San Francisco Zoo in 1971, and Dr. Francine Patterson began teaching the gorilla sign language that became part of a Stanford University project in 1974.

The foundation says Koko’s capacity for language and empathy opened the minds and hearts of millions.

Koko appeared in many documentaries and twice in National Geographic. The gorilla’s 1978 cover featured a photo that the animal had taken of herself in a mirror.

In 2016, Koko starred in an Instagram video with Flea, bassist for the Red Hot Chili Peppers.

The foundation says it will honor Koko’s legacy with a sign language application featuring Koko for the benefit of gorillas and children, as well as other projects.

Koko’s full name was Hanabi-Ko — Japanese for “fireworks child.”

(© Copyright 2018 CBS Broadcasting Inc. All Rights Reserved. The Associated Press contributed to this report.)

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